Boris Johnson hopes to stand in the west London seat of Uxbridge and South Ruislip at next year's General Election.
Mr Johnson announced earlier this month he would "in all probability… try to find somewhere to stand in 2015" - and on Tuesday he submitted his application 48 hours before the deadline.
The constituency is available because Conservative MP John Randall is standing down.
The local party will pick its candidate from up to 100 hopefuls on September 12.
David Cameron has welcomed Boris Johnson's candidacyMr Johnson said he hoped to "make his case" that he was the best person to represent the constituency.
"I'm sure there will be plenty of excellent candidates and I hope very much to make my case to the association," he told the Evening Standard.
Prime Minister David Cameron was among the first to welcome news of Mr Johnson's candidacy when it emerged on August 6.
He tweeted: "Great news that Boris plans to stand at next year's general election - I've always said I want my star players on the pitch."
Johnson was MP for Henley between 2001 and 2008There has been speculation Mr Johnson will eventually challenge for the Tory leadership.
Sky's Political Correspondent Anushka Asthana said: "David Cameron has welcomed him and thinks he could help the Conservatives win the 2015 General Election, but he might be a little worried about what happens after that."
Uxbridge and South Ruislip had a Conservative majority of more than 11,000 at the last election.
Mr Johnson has said that if he is elected to the Commons next May he will still serve out his term as Mayor, which ends in 2016.
The constituency's Labour candidate, Chris Summers, has claimed the high-profile Tory could face a backlash there because of his proposals to close the nearby Heathrow Airport and replace it with a new one in the Thames Estuary.
Boris Johnson's views on Heathrow have upset some in west LondonBack Heathrow's campaign co-ordinator Rob Gray said: "Boris Johnson's selfish plan to stand in the same borough as Heathrow is a slap in the face to tens of thousands of people in west London.
"These people rely on Britain's hub airport and the Mayor wants to close it which would result in more than 100,000 people losing their jobs, including many in Uxbridge and South Ruislip."
Mr Johnson was MP for Henley between 2001 and 2008 and served in the shadow cabinet.
News of his application comes on a day when Nigel Farage will battle with other UKIP members hoping to stand as an MP in the South Thanet constituency.
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